In a finish few saw coming when the tournament began five weeks ago, the lowest-seeded team in the field clinched the continental title with a header in the 119th minute of extra time.
The goal, scored by a 22-year-old substitute making just his third appearance of the tournament, sent the traveling section of supporters into scenes that will likely be replayed for years.
The winning side's manager, who took the job on an interim basis after the previous coach was dismissed midseason, credited a change in defensive shape for the team's improbable run through the knockout stages.
For the runners-up, who had been favorites heading into the final, the defeat extends a difficult stretch that has now seen the club fall short in three major finals over the past four years.